Born in Syracuse, NY, USA - April 21, 1939.
Raised in Huntington, NY and Cortland, NY
BA (Zoology, High Honors) - Swarthmore College, 1961.
PhD (Anatomy) - University of Chicago, 1966.
MD (Honors) - University of Chicago, 1967.
Post-doc - NIH (NINDS), 1967-1972.
Faculty, UCLA/Jules Stein Eye Institute, 1972-1980.
Professor, University of Calgary, 1980-present.
Personal interests: music - performed on flute, saxophone, voice.
Favorite activity: Mentoring students.